

Shoji Kawamura (PhD, Professor)


Muhammad Shoaib Akhtar
Project Researcher, 2025.1-
Min Hou
Project Researcher 2024.10-


PhD program
WANG Yizhen
2025.4-
2023.4-2025.3, MS
Zhixin Wu
2024.10-
2022.10-2024.9, MS
Dongyue Wang
2023.4-
2021.4-2023.3, MS
Qinyuan Ji
2021.4-
2019.4-2021.3, MS
Master program


Ziqiao Lin
2022.10-2024.9, MS
Research Assistant 2024.10-2025.2
Siyuan Chen
2020.4-2022.3, MS
Satoko Nakashima
2020.4-2022.3, MS
Zhen Ren
2020.4-2022.3, MS
Muhammad Shoaib Akhtar
2019.4-2022.3, PhD
Thesis title: Genetic variation of olfactory receptor multigene family in humans
Postdoctoral Fellow, 2022.4-2022.9
Shu Sun
2019.4-2021.3, MS
Min Hou
2018.4-2020.3, MS
2020.4-2024.9, PhD
Thesis title: Evolutionary diversification of olfactory and taste receptor gene family
compositions between omnivorous cercopithecine and highly folivorous colobine primates
Akari Kodera
2015.10-2017.9
Mizuki Endo
2015.4-2017.3, MS
Masahiro Hayashi
2015.4-2017.3, MS
Ayaka Kubota
2015.4-2017.3, MS
Takumi Naoi
2015.4-2017.3, MS
Naoki Tanaka
2015.4-2017.3, MS
Yuriko Kuriyama
2012.4-2014.3, MS
Masahiro Otoh
2012.4-2014.3, MS
Daisuke Kasai
2011.4-2013.3, MS
Masaaki Noguchi
2011.4-2016.3
Masanori Aso
2010.4-2012.3, MS
Naoya Ishizaki
2010.4-2012.3, MS
Kodama Sakurai
2010.4-2012.3, MS
Ryuichi Ashino
2009.4-2011.3, MS
2011.4-2015.9, PhD program
2015.10-2021.3, Research Assistant
Yuka Matsushita
2009.4-2011.3, MS
2011.4-2015.9, PhD program
2019.9, PhD
Thesis title: An evolutionary reconsideration of primate trichromacy through the study of L/M opsin genes
Visiting Researcher 2022.4-2025.3 (Project Researcher 2019.10-2022.3, Research
Assistant 2015.10-2019.9)
The Best Poster Award The 3rd Joint Symposium on Integrated Biosciences between Zhejiang University and the University
of Tokyo, 2019
Yukiko Fukuyo
2008.4-2009.3, Undergraduate study
2009.4-2012.3, MS
Kae Koganebuchi
2008.4-2010.3, MS
Makiko Nakata
2008.4-2010.3, MS
Akiko Sato
2008.4-2010.3, MS
Excellent Paper Award 2008 in Department of Integrated Biosciences
Naoko Takishima
2008.4-2010.3, MS
2010.4-2010.8, Laboratory Assistant
Shigeki Nakagome
2007.4-2010.3, PhD
Thesis title: An evolutionary study of genome regions associated with Crohn's disease
based on the "Out of Africa" human dispersal model
Research Fellow of JSPS 2009.4 - 2010.3
Takafumi Katsumura
2007.4-2009.3, MS
2009.4-2012.3, PhD
Thesis title: Establishing an Experimental System for the Functional Analysis of Human
Genetic Polymorphisms Using Wild Medaka (Oryzias latipes) Populations
Excellent Doctoral Thesis Award 2012
Research Fellow of JSPS 2010.4 - 2012.3
Yusuke Shirai
2007.4-2009.3, MS
Tomohide Hiwatashi
2006.4-2010.3, PhD
Thesis title: Variation of color vision and L-M opsin genes in primates
Hirotaka Matsukusa
2006.4-2008.3, MS
Yugo Okabe
2006.4-2008.3, MS
Excellent Paper Award 2006 in Department of Integrated Biosciences
Ryoko Masuda
2005.4-2007.3, MS
Norihiro Ozawa
2005.4-2007.3, MS
Sadayoshi Seno
2005.4-2007.3, MS
Tomohiro Hosoya
2004.4-2006.3, MS
Hiroko Okuda
2004.4-2006.3, MS
Toko Tsutsui
2004.4-2006.3, MS
Taro Tsujimura
2003.4-2005.3, MS
2005.4-2008.3, PhD
Thesis title: Regulatory mechanisms for the coordinated expression of duplicated green-sensitive
and red-sensitive opsin genes in zebrafish and their evolution
Excellent Doctoral Thesis Award 2008
Dean Prize, Graduate School of Frontier Sciences (Doctoral Course) 2008
Research Fellow of the Japan Society for the promotion of Science (JSPS) 2007.4 - 2009.3
Nominated for Walter M. Fitch Student Award in Annual Meeting of the Society of Molecular Biology and Evolution (SMBE) 2008
Postdoctoral Researcher Travel Award in Annual Meeting of the Society of Molecular Biology and Evolution (SMBE) 2010
Yoshifumi Matsumoto
2002.4-2004.3, MS
2004.4-2007.3, PhD
Thesis title: Gene duplication and spectral differentiation of visual opsins among closely
related medaka (Oryzias) species as a model to understand diverse evolution of fish visual systems
Research Fellow of JSPS 2006.4 - 2008.3
Best Oral Presentation Award in 2007 Primate Society of Japan Annual Meeting
Kenji Nagao
2002.4-2004.3, MS
Kumiko Takeshita
2002.4-2004.3, MS
Akito Chinen
2001.4-2004.3, PhD
Thesis title: A study of the spectral tuning and the cell type specific expressional regulation of zebrafish visual pigments for elucidation of the evolution of color vision
Excellent Doctoral Thesis Award 2004
Research Fellow of JSPS 2002.4-2004.3
Chihiro Hiramatsu
2001.4-2003.6, MS
2003.4-2006.6, PhD
Thesis title: Adaptive significance of color vision polymorphism in natural populations
of New World monkeys
Excellent Doctoral Thesis Award 2006
Research Fellow of JSPS 2003.4 - 2006.3
Takashima Prize 2010 of Primate Society of Japan
Naoya Kubotera
2001.4-2003.3,MS
Masaki Takechi
2001.4-2003.9, PhD
Excellent Paper Award 2001 in Department of Integrated Biosciences
2003.10-2006.9, PhD
Thesis title: Analysis of expression pattern of multiple red and green subtype opsin
genes and identification of cis-regulatory regions of blue and UV type opsin genes in zebrafish
Research Fellow of JSPS 2004.4-2006.9
Yukihiro Yamada
2001.4-2003.3,MS
Satoshi Kasagi
2000.4-2002.3, MS
2002.4-2007.3 in PhD program
2007.4-2011.11, Laboratory Assistant
Naomi Takenaka
2000.4-2002.3, MS
Takanori Hamaoka
1999.4-2001.3, MS
2001.4-2004.3, PhD iwork done in RIKEN Brain Science Institute Hitoshi
Okomoto Lab)
Thesis title: Visualization of visual pathways in zebrafish
Project Faculties
San San Hmwe
Project Professor, 2018.10.12-2019.5.31 (PhD, Professor, University of Yangon)
James P. Higham
Project Associate Professor, 2018.6.18-7.18 (PhD, Associate Professor, New York University)
Amanda D. Melin
Project Assistant Professor, 2018.6.18-7.18 (PhD, Assistant Professor, Canada Research Chair, University of Calgary)
Postdoctoral Fellows
Mari Nishikawa
Visiting Researcher, 2021.4-2023.3
JSPS Postdoctoral Fellow, 2018.4-2021.3
Eva Wikberg
JSPS Postdoctoral Fellow, 2012.11-2014.10
Nang Wah Wah Min
Visiting Researcher, 2010.4-2012.3
Foreign Cooperative Researchers
Miriam Pablo-Rodrigues
Universidad Veracruzana, 2013.9.2-12.1
Fernando Campos
University of Calgary, 2008.9.16-2008.12.15, 2012.11-2014.10
Amanda D. Melin
University of Calgary, 2006.2-2006.3, 2008.3-2008.4,
2010.9-2010.11
Mackenzie L. Bergstrom
University of Calgary, 2010.9-2010.11
Internship Studens
Yassmine El Yandouzi
Lyon-Tokyo Exchange program: University Claude Bernard Lyon 1, France, 2021.2.1-7.31 (Online)
Enya OfConnell-Hussey
UTSIP: Trinity College Dublin, University of Dublin, Ireland, 2019.6.23-8.3
Ruchi Shah
UTSIP: Motilal Nehru National Institute of Technology Allahabad, India, 2018.6.3-7.14
Yuchen Ma
Zhejiang University - University of Tokyo Summer Internship Program: Zhejiang University, China, 2016.6.3-8.6
Alison Paige Miller
UTSIP: Adelphi University, USA, 2015.6.29-8.7
Lauren Jacquier
Lyon-Tokyo Exchange program: Ecole Normale Superior (ENS) Lyon, France, 2015.2.23-5.29
Mehmet Nizamoglu
UTSIP: Middle East Technical University, Turkey, 2014.6.24-8.7
Kaile Zhou
UTSIP: Zhejiang University, China, 2014.6.24-8.7
Atsuko Saito
Visiting Master Program Student (University of Tokyo, Graduate School of
Arts & Sciences, Department of Life Sciences, Toshikazu Hasegawa Lab),
2000.4-2000.7
